The Human-AI Substitution Principle: When will you be replaced by AI in your organization?

Artificial Intelligence (AI) is rapidly transforming organizations, raising a fundamental organizational and economic question: when will a human employee be replaced by AI? We present an analytical model for studying Human--AI Task Allocation (HAT) in hierarchical organizations. A central feature of the HAT model is that it formally encodes the economic asymmetry between human skill acquisition and AI capability scaling. The HAT model allows us to derive how risk-adjusted costs, skills, organizational depth, deployment scale, strategic adaptation, and risk jointly determine when, where, why, and under what structural conditions human--AI replacement occurs. A key result is the Human--AI Substitution Principle, which provides a precise condition --- grounded in the formal asymmetry assumption --- under which AI replaces human labor. Building on this result, we show that AI adoption can produce abrupt workforce transitions, hybrid human--AI organizations, including cases where risk heterogeneity sustains human and AI roles without requiring a minimum-human-fraction constraint, and flatter managerial hierarchies with wider spans of control. The HAT model identifies structural conditions under which middle-management roles exhibit elevated vulnerability to automation, and shows that the vulnerability of highly skilled workers depends on a skill threshold shaped by organizational depth, baseline costs, and risk differentials. More broadly, the paper connects automation economics, organizational design, AI governance, and workforce planning into a unified theory of AI-driven organizational transformation.